Copaxa of Panama
Inspried by and dedicated to Terry Stoddard, February 19, 2013

Copaxa of Panama

Copaxa syntheratoides male, 104mm, Mount Totumas Lodge,
Amistad National Park, Chiriqui, Panama,
August 12, 2012, 1900m, courtesy of Terry Stoddard, id by Bill Oehlke.

The Copaxa belong to the Saturniini tribe, Boisduval, 1837 of the Subfamily Saturniinae, Boisduval, [1837] 1834 in the Saturniidae Family. These moths fly from Mexico into northern South America (Brazil). In most cases there appear to be at least three broods annually and both males and females come in to lights, except for the diurnal males at high altitude.

Larvae spin cocoons.

Males of several species exhibit different color morphs within the species.

Lemaire divided them into five main groups (expanded to eight by Brechlin & Meister, 2010) based on the following characters:

Decrescens Group, based on a single small (non-ringed) discal spot on each wing and a straight forewing submarginal line:
Ia) decrescens, andescens, marona, niepelti=ignescens, troetschi, yungescens, koenigi, yungaskeonigoides, ?cineracea?,
Ib) rufinans, pararufinans, rudloffi, moinieri, curvilinea, appolinairei, winbrechlini, wolfei.


II) Multifenestrata Group, based on several small discal spots on the forewing and usually on hindwing (except for expandens and andensis) and a straight forewing submarginal line:
m. multifenestrata, rufotincta, rufa, escalantei, satellita, yungallita, cineracea, denhezi, expandens, andensis, andorientalis.

III) Expandens Group:
expandens, expandens brunneocaeca, yungaspandens, cuscoexpandens, satipoexpandens,

sophronia, syntheratoides, denda, trottierorum, litensis, novocineracea,

andensis, pascoandensis, cuscoandensis.

IV) Canella Group, based on typical saturniine eyespot on each wing and straight forewing submarginal line:
IVa) Canella Subgroup:
canella, flavina, flavobrunnea, joinvillea, miranda, simson.

IVb) lavendera subgroup:
lavendera, lavenderoguatemalensis, lavenderohidalgensis, lavenderojaliscensis, parvohidalgensis; not represented in Panama

V) Sapatoza Group: High Altitude Group, based on a typical saturniine eyespot or kidney-shaped translucent discal spot on each wing and wavy forewing submarginal line:
intermediata, orientalis, semioculata, sapatoza, bella, lunula, bachuea, anikae, sinjaevi, chrisbrechlinae, herbuloti, simoni, carolinae; not represented in Panama

VI) Medea Group:
medea, ockendeni; not represented in Panama

VII) Cydippe Group:
cydippe, haxairei, evelinae; not represented in Panama

VIII) Copaxoides Group, based on typical saturniine eyespot on each wing and wavy forewing submarginal line:
copaxoides, muellerana, mannana; not represented in Panama

Copaxa syntheratoides male (pink form), 111mm, Mount Totumas Lodge,
Amistad National Park, Chiriqui, Panama,
August 12, 2012, 1900m, courtesy of Terry Stoddard, id by Bill Oehlke.

Please note: The advent of DNA barcoding has resulted in many new descriptions. In many cases the "new" species are quite similar to existing species. Do not be surprised if even more refined testing or revisions of thresholds of difference in the future results in some synonymies or even more species/subspecies. Subsequent rearing may or may not indicate differences in larval appearance. It will be interesting to see how this all pans out.

Decrescens Group 1A
Single, small, non-ringed, hyaline spot on each wing
straight pm line
Copaxa decrescens, the type for this group, is limited to southeastern Brazil and surrounding areas, and would not be found in Panama.

Copaxa ignescens is similar to decrescens and may or may not fly in southern Panama. It has not been documented there as yet, but has been taken in western Colombia.

Look-a-like moths in Panama would probably be Copaxa troetschi.

Decrescens Group 1A
Single, small, non-ringed, hyaline spot on each wing
straight pm line
ignescens
Brechlin & Meister image does not show strong red presence; rather an almost uniform beigey-brown
not confirmed in Panama

Decrescens Group 1A
Single, small, non-ringed, hyaline spot on each wing
straight pm line

troetschi
Note paler, pinkish fw basal area continuing into lower quarter of median area to the thin, scalloped line

Decrescens Group 1B
Single, small, non-ringed, hyaline spot on each wing
heavy pink suffusion in pm area
rufinans
falcate forewing apex
straight transverse line

Decrescens Group 1B
Single, small, non-ringed, hyaline spot on each wing
heavy pink suffusion in pm area
moinieri
male fwl: 60-62mm
relatively small fw cell mark
uniform, pale brown gc

Decrescens Group 1B
Single, small, non-ringed, hyaline spot on each wing
heavy pink suffusion in pm area

curvilinea
very falcate forewing apex
curved transverse line

Multifenestrata Group 2
fw: several small discal spots on forewing usually on hindwing
straight forewing submarginal line
multifenestrata

Multifenestrata Group 2
fw: several small discal spots on forewing usually on hindwing
straight forewing submarginal line
conlani

Multifenestrata Group 2
fw: several small discal spots on forewing usually on hindwing
straight forewing submarginal line
rufotincta
This species is confirmed for Panama (possibly a recording error??)
I do not think the specimen depicted above is representative of rufotincta.





Multifenestrata Group 2
fw: several small discal spots on forewing usually on hindwing
straight forewing submarginal line
australoescalantei
Brechlin and Meister, 2010, report C. australoescalantei from Nicaragua; Costa Rica and Guatemala at 1100-1540m, refering to it being slightly smaller than the Mexican C. escalantei
C. australoescalantei might fly in Panama, or it might be replaced there by C. rufotincta.

Multifenestrata Group 2
fw: several small discal spots on forewing usually on hindwing
straight forewing submarginal line



escalantei
I am not sure if escalantei is limited in its range to Mexico and Belize or whether it also flies further south and might be sympatric with australoescalantei.




Expandens Group 3
syntheratoides
This species is very variable with regard to colouration and patterning, but the forewing apex is generally much less pointed than in other Panamanian species.
The single small hyaline spot on each wings seems a consistent character. The same hyaline spots are considerably larger in the females, all of which seem to be yellow with varying amounts of brown markings.

Expandens Group 3
syntheratoides
pink form
This species is very variable with regard to colouration and patterning, but the forewing apex is generally much less pointed than in other Panamanian species.
The single small hyaline spot on each wings seems a consistent character. The same hyaline spots are considerably larger in the females, all of which seem to be yellow with varying amounts of brown markings.

Canella Group 4 based on typical saturniine eyespot on each wing and straight forewing submarginal line
Canella Subgroup 4A
simson
This is the only Panamanian Copaxa species that have a larger, single circular ocellus on each wing. Females tend to be yellow beige, instead of the orange found in males.




Unconfirmed Possibilities

rudloffi
male fwl: 48-54mm; 860m
Nicaragua, Costa Rica
similar to moinieri, but smaller
red and brown forms
more contrast than monieri

Neither C. rudloffi nor C. trottierorum have been reported in Panama, but both have been documented in Costa Rica so one or the other or both may turn up in Panama.

C. rudloffi, in the Decrescens Group 1B, is most similar to C. moinieri, but moinieri is considerably larger with less contrast.

C. trottierorum, in the Expandens Group 3, is quite distinct with the white suffusions at apex and outside the submarginal line.

trottierorum
reported in eastern Costa Rica
strong presence of white suffusion near forewing apex and outside submarginal line




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